Denitrification has a wide range of factors that control its complex regulatory network, one of which is the low concentration of electron donors. In this case the absence of available electrons causes high concentrations of cytotoxic gases (NO and/or N2O) for denitrifying bacteria. The individual adaptive behavior to environmental conditions through different behavior-rules is one of the most powerful aspects of microbial individual-based models.
Araujo, P., Gras, A., Ginovart, M. Mass degradation to reduce cytotoxic products as an individual behavior-rule embedded in a microbial model for the study of the denitrification process. A: International Conference on Environmental, Industrial and Applied Microbiology. "BioMicroWorld 2015 - Book of Abstracts". 2015, p. 653.